When Variation Doesn’t Become a Boundary: A Putative Mixed Subspecies Pairing in Long-tailed Tits
A familiar presence in Central European woodlands, the Long-tailed Tit Aegithalos caudatus europaeus carries the darker head pattern that defines much of the region’s breeding population—an anchor against which rarer, paler forms reveal themselves. © Alexis Lours (CC-BY)

When Variation Doesn’t Become a Boundary: A Putative Mixed Subspecies Pairing in Long-tailed Tits

A pair of Long-tailed Tits reveals how subspecies boundaries, so clear on paper, begin to soften in the field when variation and behaviour intersect.

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